Honey bees, Apis mellifera, occur naturally over vast and varied geographical areas, extending from Scandinavia in the north to the Cape of Good Hope in the south, and from Dakar in the west to Oman in the east. Different populations are adapted to a very wide range of climatic conditions (e.g., Ruttner et al., 1978). Apis mellifera colonies are found from sea level to 1,000 m above sea level (a.s.l.) in temperate zones and from sea level to 3,700 m a.s.l. in the tropics. They also survive in the hot and arid zones of Oman at 200 m a.s.l. (Dutton et al., 1981).